Editing Cat (section)

== Behavior ==
See also: Cat behavior

Outdoor cats are active both day and night, although they tend to be slightly more active at night. Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity is quite flexible and varied but being low-light predators, they are generally crepuscular, which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent.

Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.

A common misconception is that a cat's behavioral and personality traits correspond to its coat color. These traits instead depend on a complex interplay between genetic and environmental factors.

=== Sociability ===
The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling, and, if that does not work, by short and violent, noisy attacks. Though cats do not have a social survival strategy or herd behavior, they always hunt alone.

Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.

Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression, there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, pet, human or other being.

Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means of social bonding.

=== Communication ===
Main article: Cat communication

Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms; a raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent. Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.

Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.

The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).

Domestic cats observed in rescue facilities have 276 morphologically distinct facial expressions based on 26 facial movements; each facial expression corresponds to different social functions that are likely influenced by domestication. Facial expressions have helped researchers detect pain in cats. The feline grimace scale's five criteria ear position, orbital tightening, muzzle tension, whisker change, and head position indicated the presence of acute pain in cats.

=== Grooming ===
Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 0.5 millimeter long, called lingual papillae, which contain keratin making them rigid. The papillae act like a hairbrush, and some cats, particularly long-haired cats, occasionally regurgitate sausage-shaped 2–3 cm (0.79–1.18 in) long hairballs of fur that have collected in their stomachs from grooming. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.

=== Fighting ===
Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.

When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways, and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering slaps to the face and body with the forepaws, as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their hind legs.

Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe, and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections from scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of the feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.

=== Hunting and feeding ===
See also: Cat food

The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking, they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.

Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell, and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.

Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Other common small creatures, such as lizards and snakes, may also become prey. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds. Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.

Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (''Erithacus rubecula'') and dunnocks (''Prunella modularis'') in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes, which prey on cats and other small predators, reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.

=== Play ===
Main article: Cat play and toys
[[/wiki/File:Play_fight_between_cats.webmhd.webm|thumb|Play fight between kittens aged 14 weeks]]
Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, both with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and it might also reduce the fear that they associate with launching attacks on other animals.

Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death.

=== Reproduction ===
See also: Kitten

The cat secretes and perceives pheromones. Female cats, called ''queens'', are polyestrous with several estrus cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August in northern temperate zones and throughout the year in equatorial regions.

Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (0.039 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.

After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.

The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.

Based on a study of 2,300 free-ranging queens conducted from May 1998 and October 2000, they had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.

Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males, and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.